There are two types of growth that take place in plants. They are primary growth and secondary growth. The primary growth is related to the growth of apical meristems whereas the secondary growth is related to the growth of roots and stems in the lateral meristems. Secondary growth is shown by the dicotyledon plants. Due to a lack of vascular tissue, secondary growth does not occur in monocots. The wood plant tissue is formed by the secondary growth in dicot stems. This wood plant tissue is responsible for providing protection to the plant. The lenticels present in the wood material helps in the exchange of gases. We will learn about how this growth takes place and about the wood yielding plants that are useful for us. The wood obtained from the plants is also used in making paper. It also finds its uses in making furniture and shelter.
As we studied above, the secondary growth in plants is responsible for the formation of wood. The vascular bundles get arranged in a ring-like manner. They are arranged around the central pith and are conjoint and open. As they possess cambium therefore they are called open tissues. It is known as intrafascicular cambium. The cells start the process of dedifferentiation and in this way, the cambium rings are formed. These cambium rings that are formed by the cambium start dividing. From the observations, it is seen that the cambium is more active on the inner side as compared to the outer side.
The cork cambium is the main tissue that is responsible for the formation of wood. This will give us a brief idea about wood information. The girth of the stem increases due to increased activity of the vascular cambium. As the girth keeps increasing the outer cortical layers starts rupturing. So, cork cambium produces new layers that replace the damaged or ruptured layers. Cork cambium is also called phellogen. The cork cambium has another name that is stellar cambium. Phellogen is thick and has two layers. The outer one forms the cork and the inner one forms the secondary cortex. The cells of cork are compactly arranged and in the beginning, they have thin cellulose cell walls. When they mature the living part is replaced by the non-living part which is the formation of wood material. The cell walls of the cork become thick by the deposition of suberin. This chemical makes the cork or wood material impervious to water by getting deposited in the cell walls. The phelloderm is the secondary cortex. It is called so because it develops at the time of secondary growth. It is made up of thin-walled parenchymatous cells. They have cellulose cell walls and are living in nature. The periderm is the collective name given to phellogen, phellem and phelloderm. They are the protective layers of the cell. They grow when the epidermis layer is ruptured and also when the outer cortical layers are ruptured. When secondary growth in the vascular cambium takes place then only the secondary growth of the cork cambium happens. As the growth of cork cambium is continuous, the layers peripheral to phellogen are damaged and they need continuous replacement.
The wood of the tree is mainly present in its bark. The bark is made up of all the tissues that are present outside the vascular cambium. The layers that make up the bark are the periderm, primary cortex, pericycle, primary and secondary phloem. These all tissues are present outside the vascular cambium. The bark formed in the early season is known as soft bark and the bark that is formed later in the season is called hard bark. When the entire ring of the cork cambium makes a bark, it is called ring bark. In this, a complete cylinder of bark is formed.
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Figure: Wood made from the bark of the tree
They are the small openings that are formed in the bark. They are formed by a small portion of the periderm. They are produced by the activity of the phellogen. As we read above, the phellogen is the meristematic tissue that is formed during the secondary growth of plants. They are lens-shaped openings. As they are openings, they help in the exchange of gases. The exchange of gases takes place between the internal tissue of the stem and the outer atmosphere. They help in exchanging gases from the woody areas of the plant. Water is also lost from them in the form of vapours. As they help in the exchange of gases, they are also known as breathing pores.
The wood that can be used for carpentry purposes is known as timber. It is of two types: heartwood and softwood. Some common timber yielding trees are:
Teak
Sal
Sissoo
Sins
Arjun
Harir
Deodar
There are two other types of wood that are known as heartwood and sapwood. Heartwood is a modified and non-functional secondary xylem. Due to the deposition of organic compounds, the heartwood becomes resistant to microbial growths. It also becomes hard and durable in nature. It has dead elements and the cell walls are highly lignified. No water is conducted from this heartwood. It helps in providing mechanical support to the plant. The sapwood is present in the periphery region of the secondary xylem. It is actively involved in conducting water and minerals. Due to the passage of time and the addition of layers in the cork cambium, more and more sapwoods are changed to heartwoods.
1. What is wood from a botanical perspective?
In botanical terms, wood is technically the secondary xylem produced by the vascular cambium during secondary growth in the stems and roots of dicotyledonous and gymnosperm plants. It is a complex plant tissue that provides structural support to the plant and helps in the conduction of water and minerals from the roots to the leaves.
2. How is wood formed in plants during secondary growth?
Wood formation is a result of secondary growth, which increases the girth of the stem. The process is driven by the activity of a meristematic layer called the vascular cambium. This cambium is located between the xylem and phloem. It divides to form new cells, producing secondary xylem towards the inner side (the pith) and secondary phloem towards the outer side. The continuous accumulation of this secondary xylem over the years leads to the formation of the bulk of tissue we call wood.
3. What is the key difference between heartwood and sapwood?
Heartwood and sapwood are two distinct regions within the wood of a tree, differing mainly in function, location, and composition. The key differences are:
4. Why is heartwood typically darker and more resistant to decay than sapwood?
Heartwood is darker and more durable due to significant physiological and chemical changes as it ages. The living parenchymatous cells die, and the tracheary elements get plugged by tyloses (balloon-like outgrowths). Furthermore, the cells become filled with organic compounds like tannins, resins, oils, and gums. These substances make the heartwood dark, hard, and highly resistant to attacks by microbes and insects, thus enhancing its durability.
5. What are lenticels and what is their importance in a woody stem?
Lenticels are porous, lens-shaped openings found on the bark of woody stems. They are formed by the phellogen (cork cambium), which produces loosely arranged parenchyma cells that rupture the epidermis. Their primary importance is to facilitate the exchange of gases (like oxygen and carbon dioxide) between the internal living tissues of the stem and the outside atmosphere, effectively acting as 'breathing pores' for the stem.
6. Is bark the same as wood? Explain the difference.
No, bark and wood are not the same; they are distinct parts of a woody stem. Wood is specifically the secondary xylem located inside the vascular cambium. Bark, on the other hand, is a non-technical term that refers to all tissues located outside the vascular cambium. This includes the secondary phloem, primary cortex, and the periderm (which consists of phellogen, phellem, and phelloderm).
7. What are some common examples of woody plants that are important commercially?
Many woody plants, or trees, are commercially valuable for their timber. Some well-known examples based on their uses include: