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Introduction to Anther

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In botany, it is as important to study the parts and functions of the flower as it is that of plants. The anther is a part of a flower. Let’s start by learning the parts of a flower, pistil, stamen, receptacle, peduncle and sepal. To define anther, it is crucial to understand the reproductive parts of the flower. The female reproductive part of a flower is the pistil, composed of stigma, style and ovary. And the male reproductive part of the flower is called the stamen. The anther is the part of a flower’s male reproductive system stamen. The male structure stamen is composed of a filament that is a long tube with an oval-shaped pollen-producing structure attached to the top of it called the anther. The anther is solely responsible for producing and bearing pollen that serves the purpose of fertilization in flowering plants. Only upon pollen production by the anther does the pollination commences.  


Male Reproductive Part of the Flower

The male part of the reproductive system of the flower is called Androecium that is individually segregated as the stamen. The stamens are elongated filaments raised in the middle of the flower in a circular ring. There are at least five to six stamens in a flower. The ratio of the number of stamens is the same as that of the petals but sometimes it varies depending on varieties of flowers and their species. Anther of a flower and filament are the two parts of the stamen that surround the pistil. Anther meaning and role can be particularly understood better when you look at the diagram below that suggests the location of the anther. It also depicts the other parts very clearly for a clearer understanding and one can refer to when reading about anther. 

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Parts of a Flower

As understood from the above diagram, stamens are separated but all connected and collectively the stamens are called androecium. A filament of the stamen very diligently holds the anther on top and facilitates its functions. The filament holding the anther is attached to the petal, and the nature of the stamen solely depends on the type of the plants and flowers. The plants that reproduce through self-pollination which is via the transfer of the pollen from the anther to the stigma of the pistil to reach the ovary of the same flower of the same plant has the stamen bent towards the pistil. In the case of plant reproduction via cross-pollination that is the stigma receives the pollen from flowers of other plants via various means like insects and wind the stamen is usually bent away from the pistil. 


Anther and Stigma 

Anther of a flower is often confused with stigma even though their functions in the flower serve different purposes. Let’s take a look at what the anther contains versus the stigma. Anther and stigma are the male and female reproductive parts of a flower respectively. 


Anther

Stigma

The anther is part of the male reproductive system, androecium in a flower.

The stigma is part of the female reproductive system, gynoecium in a flower.

The anther is connected to the distal end of the filament and is also the main attraction for the pollinators like insects and animals.

The stigma is also located in the distal end of the pistil and it definitely plays an important role via its texture and shape to hold and transport the grain. 

The anther has an oval-shaped structure and can be of one lobe or two.

The stigma of a flower can be long, slender, feathery or even globe-shaped depending on the type of species.

The anther is at the top and held by a long tube called the filament.

The stigma lies on the top and is held by a tube called the style that connects to the ovary.

The anther and filament together form the stamen the individual part of androecium.

The stigma, style and ovary together form the pistil and this is the composition of the gynoecium.

The anther contains, bears and produces pollen grains or pollens that are like the sperm of a flower for fertilization. 

The stigma on the other hand receives this pollen grain and the compatible ones reach the ovary allowing germination and reproduction.

Since the anther of a flower contains pollen grains it is grainy in texture and feels. 

The stigma is sticky since its function is to receive, retain and facilitate the movement of pollen for germination.


Structure and Function of the Anther

The function of the anther is to produce and provide the pollen, the pollen containing anther acts as the sperm that is necessary for reproduction. The anther functions in attracting pollinators, such as bumblebees and other insects to the flower who via sucking on the nectar does aid in both self-pollination and cross-pollination. And reproduction would be impossible without this crucial part of the flower's anatomy that is the anther. 

For the understanding of pollen production, it is important to understand the structure of the pollen-producing oval structured anther.  

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Transverse Section of the Anther of a Flower

The anther which is the fertile part of the stamen usually has two lobes or thecas connected by a connective. Each lobe contains two pollen chambers longitudinally placed. Each chamber filled with pollen or pollen grains is also called microspores and hence these chambers are also called microsporangium. 

Four layers in total surround the pollen sac or pollen chambers, and they are namely, Epidermis, the inner epidermis or endothecium, the middle layer and the tapetum. 

  1. Epidermis:- it is the outermost layer and is a single layer that is of thickened fibrous matter whose function is to protect the pollen sacs.

  2. Endothecium or the inner epidermis is a single layer of radially elongated cells that develops cellulose with pectin and lignin fibrous thickening around the sac which holds it together and prevents it from splitting or bursting open.

  3. The middle layer is a thin-walled cell of three to four layers situated just below the endothecium nourishes the microspore for facilitating pollen production. 

  4. The fourth layer that lies closest to the pollen sac is nutritive and provides necessary nutrition for pollen division and pollen generation.

The anther with two thecas are called dithecous anther and these are very common. Dithecous means two thecas or lobes it can also be referred to as bilobed. Some examples of bilobed or dithecous anther containing flowers are Solanum and Crotalaria. Some flowers of the plants, however, only have one lobe or theca that produces pollen. Such ones are called monothecous anther. Examples of a monothecous anther are Hibiscus.


Fun Facts about Anther

  1. Since the anther’s main function is to aid reproduction via pollination, the base of the stamen consists of nectaries that attract the pollinators like insects, mainly bees like bumblebees to carry out this function.

  2. The nectaries provide food to these pollinators as a reward from nature to carry out pollination.

  3. The pollination process via the anther is also carried out by water, wind, animals and birds.

  4. The microspore containing pollen sac that produces pollen is called microsporangium or pollen mother cells.

FAQ (Frequently Asked Questions)

Q1. Is the Anther of a Flower Male or Female?

Ans. The anther of a flower is a part of the male reproductive system stamen. It is held above by the long tube called the filament. The anther of a flower is grainy as it produces and bears the pollen grains. The pollen grains transfer to the stigma of the flower to reach the ovary and fertilizes for reproduction to start. The anther that produces pollen in a way acts like the sperm so can be considered a part of the male anatomical reproductive system of a flower. 

Q2. What is the Main Role of the Anther?

Ans. The anther can be dithecous with two lobes of pollen sac or monothecous with only one lobe that is rarely the case. In both cases, the main role is to produce pollen for continuing the species of the plants. The anther of a flower does the main role of providing pollen grains to facilitate reproduction, without which there would be no more plants and the growth would cease. The anther of the flower in itself carries the main role of pollination via pollen production. The most important part for the reproduction to be proper is for pollen grains to be of good quality and the anther plays a major role in its nourishment as well. 

Q3. What is the Filament in a Flower?

Ans. The filament is the long and slender stalk or tube in the stamen that is the male reproductive part of the flower. The proximal end of the filament of the stamen is attached to the base of the flower thalamus or the petals of the flower. And the filament is raised wherein the distal end is attached to the anther. The filament carries the anther which bears and produces the pollen. 

 Q4. Does Filament Have Any Role in the Reproduction of Flowering Plants?

Ans. The role of the filament is indeed crucial for reproduction. Though the reproduction of the flowering plants has to occur only with the transfer of pollen, it is the filament that makes this transfer possible. Firstly, it provides the site of pollen development for the anther and continuously transmits nutrients and water to the pollen for proper growth. At the same time, the nectaries at the base attract pollinators. And the filament positions the anther in such a way that dispersal of pollen is possible for either self-pollination or cross-pollination.